Ray Romano

Manny (voice)

Josh Hamilton

Dodo / Aardvark (voice)

Chris Wedge

Dodo / Scrat (voice)

Denny Dillon

Glyptodon (voice)

Mitzi McCall

Glyptodont (voice)

Tara Strong

Baby Moeritherium (voice)

Darin De Paul

Additional Voices (voice) (uncredited)

Dann Fink

Dodo (voice) (uncredited)

Patrick Pinney

Various Mammals (uncredited)

Jane Krakowski

Rachel (voice)

Lorri Bagley

Jennifer (voice)

John Leguizamo

Sid (voice)

Denis Leary

Diego (voice)

Goran Visnjic

Soto (voice)

Jack Black

Zeke (voice)

Cedric the Entertainer

Carl (voice)

Stephen Root

Frank / Start (voice)

Diedrich Bader

Oscar (voice)

Alan Tudyk

Lenny / Oscar / Dab (voice)

Phil Proctor

Various Mammals (uncredited)