Ray Romano

Manny (voice)

Ester Dean

Female Siren (voice)

Eddie « Piolin » Sotelo

Uncle Fungus (voice)

Joy Behar

Eunice (voice)

Heather Morris

Katie (voice)

Chris Wedge

Scrat (voice)

Marieve Herington

Additional Voices (voice)

Simon Pegg

Buck (voice)

Patrick Stewart

Ariscratle (voice)

Edita Brychta

Additional Voices (voice)

Dennis T. Carnegie

Additional Voices (voice)

Aimée Castle

Additional Voices (voice)

Jim Conroy

Additional Voices (voice)

Jason Harris

Additional Voices (voice)

Sondra James

Additional Voices (voice)

Tara Sands

Additional Voices (voice)

Alan Tudyk

Milton / Hunky Siren (voice)

Rebel Wilson

Raz (voice)

Nicki Minaj

Steffie (voice)

John Leguizamo

Sid (voice)